A 31-year-old woman arrested by detectives investigating a hate campaign waged against the owners of a guinea pig breeding farm has been released on police bail.

The woman was detained at her home in Gloucester yesterday morning by officers investigating the theft of a pensioner's body from a graveyard in Yoxall, Stafford-shire, in 2004, and freed pending further inquiries.

Police said she was arrested on suspicion of conspiracy to blackmail David Hall and Partners - which operated Darley Oaks Farm in New-church, Staffordshire, breeding animals for medical research - and others.

Her detention follows the arrest on Tuesday of a 24-year-old Wolverhampton woman who was taken into custody by officers investigating the desecration and released on police bail yesterday.

In a statement, Stafford-shire Police said the 31-year-old woman was arrested at around 7am following a preplanned operation and taken for questioning at a police station in the county.

A force spokesman said: "She was arrested on suspicion of conspiracy to blackmail David Hall and Partners and others."

Four other people have been charged with conspiracy to blackmail the farm owners and are due to appear at Nottingham Crown Court on Monday for a plea and directions hearing.

Guinea pig breeding operations at Darley Oaks Farm ceased last month following a six-year campaign of intimidation by animal rights extremists.

The Hall family announced the closure on January 20 and said they hoped the decision would prompt grave-robbers to return the body of a relative.

The remains of 82-year-old Gladys Hammond, the mother-in-law of Christopher Hall, who co-owns the farm, were stolen from St Peter's Church in October 2004.